Fossil Record 24(2): 455-465, doi: 10.5194/fr-24-455-2022
Ingensalinae subfam. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea: Inoderbidae), a new planthopper subfamily from mid-Cretaceous Kachin amber from Myanmar
expand article infoCihang Luo, Zhishun Song§, Xiaojing Liu|, Tian Jiang, Edmund A. Jarzembowski, Jacek Szwedo#
‡ State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China§ Institute of Insect Resources and Biodiversity, School of Life Sciences, Chemistry & Chemical Engineering, Jiangsu Second Normal University, Nanjing 210013, China| School of GeoSciences, Yangtze University, Wuhan, Hubei 430100, China¶ State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences (Beijing), Beijing 100083, China# Laboratory of Evolutionary Entomology and Museum of Amber Inclusions, Department of Invertebrate Zoology and Parasitology, Faculty of Biology, University of Gdańsk, 59 Wita Stwosza St., 80-309 Gdańsk, Poland
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The second genus and species of recently establishedplanthopper family Inoderbidae, Ingensala xiai gen. et sp. nov., is described based on awell-preserved specimen from mid-Cretaceous Kachin (Burmese) amber, and itcan be definitely attributed to Inoderbidae mainly based on its headstructure, pronotum, and mesonotum without median and lateral carinae andtegmen venation. Ingensala gen. nov. is superficially similar to Eofulgoridium regarding itsvenation pattern, rather than to the Inoderbidae type genus Inoderbe, and furtherconfirmed that Inoderbidae might descend from the Jurassic planthopper familyFulgoridiidae. The early fork of CuA and the stem CuA bearing many branchesalso can be found in Jurassic Qiyangiricaniidae and Eocene Weiwoboidae.Ingensala gen. nov. also superficially resembles some Tropiduchidae: Tropiduchinae.The new genus differs from the type genus Inoderbe to a large extent according toits wide head, frons without fastigium, antennae not so elongate, thetectiform condition of wings' position in repose, large, broad andtranslucent tegmen, triangular basal cell, single CuA1, legs coveredwith short setae, and the lack of filamentous wax on body. Therefore, twonew subfamilies (Inoderbinae stat. nov. and Ingensalinae subfam. nov.) areestablished for these two genera respectively. The diversification inplanthoppers could be the result of pressure of environmental changes duringthe mid-Cretaceous, and Inoderbidae provides more information for us tounderstand the Cretaceous stage of Fulgoroidea evolution anddiversification.